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The Pillar of Autumn
Iron Skull - At the start of the level, as you awake from the cryo-stasis container, search the area behind the cryo container. It's behind some crates at the back of the room.

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Halo
Mythic Skull - As soon as the level starts and you land on Halo, you'll see a large waterfall directly ahead of you. Search the base of the waterfall and you'll find the skull around some rocks to the right.

Boom Skull - After you receive the Warthog for the first time, keep driving until you see an entrance to some underground tunnels. Before entering, look above to see some pipes. Scale the cliff edge to the right to find the skull.

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Truth and Reconciliation
Foreign Skull - Check behind one of the one-way doors from which the enemies spawn from.

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The Silent Cartographer
Famine Skull - Once you clear the first set of enemies on the beach, turn around and face the ocean. Now follow the path right and you'll see a small plateau. Search the top of it to find the skull.

Bandanna Skull - This skull is located at the end of the first facility. You'll need to grenade jump to reach it. If playing in co-op, your partner can commit suicide once you're up top so he/she can spawn next to you and get it as well. This skull is very had to get.

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Assault on the Control Room
Fog Skull - In the patrol room, before you head outside again onto the battlefield, search the vent in the center of the room, grenade jump required. Your partner should commit suicide to spawn by you if playing in co-op.

Malfunction Skull - When you reach the gorge, you'll see ladders leading down to lower platforms, jump or climb down to get the skull.

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343 Guilty Spark
Recession Skull - Check behind the first shade/tree you see in the swamp, at the start of the mission. Follow the video strictly for reference as it's hard to see.

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The Library
Black Eye Skull - From the start of the level, walk around until you reach the 3rd set of vents, just before you enter the index room for the first time. It's in the right vent; you'll need to grenade jump.

Eye Patch Skull - Avoid the index on this floor and walk around the edge of the room. It's in-between 2 crates. If you go into the index, you will miss this.

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Two Betrayals
Piata Skull - After you capture the last Banshee, fly back past the third generator room, then back to the tunnel. It's on the snow hill above the tunnel opening where you came from.

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The Maw
Grunt Birthday Party Skull - Use your Warthog to climb up on the railing where Foehammer goes down near the end of the mission. The timer stops here so it's another hard to miss skull